Genus Pachycondyla (Ponerinae)
- Pachycondyla
Smith, F. 1858a: 105. Type species: Formica crassinoda Latreille, by subsequent designation of Emery, 1901b: 42.
- Bothroponera
Mayr, G. 1862: 717. Type species: Ponera pumicosa Roger, by subsequent designation of Emery, 1901b: 42. Subgenus of Pachycondyla: Emery, 1901b: 42. Revived status as genus: Wheeler, W.M. 1918c: 299 (footnote). Synonymy:
Brown, W.L. 1973b: 179
(Bothroponera provisional junior synonym of Pachycondyla)
;
Bolton, 1995b: 22 (Bothroponera junior synonym of Pachycondyla)
.
- Megaponera
Mayr, G. 1862: 734. Type species: Formica foetens Fabricius, by monotypy. The type species, Formica foetens Fabricius, is a junior primary homonym in Formica. The replacement name is Formica analis Latreille. Synonymy:
Bolton, 1995b: 34.
- Paltothyreus
Mayr, G. 1862: 735. Type species: Formica tarsata Fabricius, by monotypy. Synonymy:
Bolton, 1995b: 41.
- Ectomomyrmex
Mayr, G. 1867a: 83
. Type species: Ectomomyrmex javanus Mayr, by subsequent designation of Bingham, 1903: 85. Subgenus of Pachycondyla: Emery, 1901b: 42. Revived status as genus: Wheeler, W.M. 1922i: 648. Synonymy:
Brown, W.L. 1973b: 180
(Ectotomomyrmex provisional junior synonym of Pachycondyla)
;
Bolton, 1995b: 28 (Ectomomyrmex junior synonym of Pachycondyla)
.
- Megaloponera
Emery, 1877a: 368. Incorrect subsequent spelling of Megaponera Mayr, 1862. Synonymy:
Bolton, 1995b: 34.
- Ophthalmopone
Forel, 1890b: cxi. Type species: Ophthalmopone berthoudi Forel, by monotypy. Synonymy:
Bolton, 1995b: 40.
- Euponera
Forel, 1891b: 126 (as subgenus of Ponera)
. Type species: Ponera (Euponera) sikorae Forel, by monotypy. Raised to genus: Emery, 1901b: 46. Synonymy:
Bolton, 1995b: 29.
- Brachyponera
Emery, 1900b: 315 (as subgenus of Euponera)
. Type species: Euponera (Brachyponera) croceicornis Emery, by monotypy. Raised to genus: Bingham, 1903: 101. Subgenus of Euponera: Emery, 1911e: 83. Maintained as subgenus: Wheeler, W.M. 1922j: 777. Raised to genus: Wilson, 1958g: 346. Also described as new by Emery 1901b: 43. Synonymy:
Snelling, 1981: 389 (Brachyponera provisional junior synonym of Pachycondyla)
;
Bolton, 1995b: 22 (Brachyponera junior synonym of Pachycondyla)
.
- Mesoponera
Emery, 1900c: 668 (as subgenus of Euponera)
. Type species: Ponera melanaria Emery, by monotypy. Raised to genus: Bingham, 1903: 99. Subgenus of Euponera: Emery, 1911e: 80. Maintinaed as subgenus: Wheeler, W.M. 1922i: 649. Revived status as genus: Wilson, 1958g: 349. Also described as new by Emery, 1901b: 43. Type species not Ponera caffraria F. Smith, unjustified subsequent designation by Emery, 1901b: 43.) Synonymy:
Brown, W.L. 1973b: 182
(Mesoponera provisional junior synonym of Pachycondyla)
;
Bolton, 1995b: 35 (Mesoponera junior synonym of Pachycondyla)
.
- Pseudoponera
Emery, 1900b: 314 (as subgenus of Pachycondyla)
. Type species: Ponera quadridentata Smith, by monotypy. Subgenus of Euponera: Forel, 1901d: 141. Raised to genus: Bingham, 1903: 91. Maintained as genus: Emery, 1911e: 86; Wheeler, W.M. 1922i: 649. Also described as new Emery, 1901b: 42. Type species not Ponera amblyops Emery, unjustifed subsequent designation by Emery, 1901b: 42. The type species, Ponera quadridentata Smith, is a junior synonym of Pachycondyla stigma Facricius. Synonymy:
Brown, W.L. 1973b: 184
(Pseudoponera provisional junior synonym of Pachycondyla)
;
Bolton, 1995b: 45 (Pseudoponera junior synonym of Pachycondyla.)
.
- Hagensia
Forel, 1901k: 333 (as subgenus of Megaloponera)
. Type species: Megaloponera (Hagensia) havilandi Forel, by monotypy. Subgenus of Euponera: Forel, 1917: 237. Raised to genus: Arnold, 1926: 202. Synonymy:
Bolton, 1995b: 30.
- Neoponera
Emery, 1901b: 43. Type species: Formica villosa Fabricius, by original designation. Synonymy:
Brown, W.L. 1973b: 183
(Neoponera provisional junior synonym of Pachycondyla)
;
Bolton, 1995b: 39 (Neoponera junior synonym of Pachycondyla)
.
- Eumecopone
Forel, 1901j: 335 (as subgenus of Neoponera)
. Type species: Neoponera (Eumecopone) agilis Forel, by monotypy. Synonymy:
Brown, W.L. 1973b: 180
(Eumecopone provisional junior synonym of Pachycondyla)
;
Bolton, 1995b: 29 (Eumecopone junior synonym of Pachycondyla)
.
- Trachymesopus
Emery, 1911e: 84 (as subgenus of Euponera)
. Type species: Formica stigma Fabricius, by original designation. Raised to genus: Wilson, 1958g: 352 (see also Brown, W.L. 1963a: 6). Synonymy:
Brown, W.L. 1973b: 185
(Trachymesopus provisional junior synonym of Pachycondyla)
;
Bolton, 1995b: 48 (Trachymesopus junior synonym of Pachycondyla)
.
- Hiphopelta
Forel, 1913a: 108. Incorrect subsequent spelling of Xiphopelta Forel, 1913. Synonymy:
Bolton, 1995b: 31.
- Xiphopelta
Forel, 1913a: 108 (as subgenus of Ponera)
. Type species: Ponera (Xiphopelta) arnoldi Forel, by monotypy. Subgenus of Euponera: Forel, 1917: 237. Synonymy:
Wheeler, W.M. 1922j: 775 (Xiphopelta junior synonym of Mesoponera)
;
Bolton, 1995b: 50 (Mesoponera junior synonym of Pachycondyla.)
.
- Trachyponera
Santschi, 1928a: 43. Incorrect subsequent spelling of Trachymesopus Emery, 1911. Synonymy:
Bolton, 1995b: 48.
- Termitopone
Wheeler, W.M. 1936e: 159. Type species: Ponera laevigata Smith, by original designation. Synonymy:
Brown, W.L. 1973b: 185
(Termitopone provisional junior synonym of Pachycondyla)
;
Bolton, 1995b: 48 (Termitopone junior synonym of Pachycondyla)
.
- Syntermitopone
Wheeler, W.M. 1936e: 169 (as subgenus of Termitopone)
. Type species: Ponera commutata Roger, by original designation. Synonymy:
Borgmeier, 1959a: 312 (Syntermitopone junior synonym of Termitopone)
;
Bolton, 1995b: 47 (Termitopone junior synonym of Pachycondyla.)
.
- Wadeura
Weber, 1939a: 102. Type species: Wadeura guianensis André, by original designation. Synonymy:
Brown, W.L. 1973b: 185
(Wadeura provisional junior synonym of Pachycondyla)
;
Bolton, 1995b: 49 (Wadeura junior synonym of Pachycondyla)
.
- Pseudoneoponera
Donisthorpe, 1943e: 439. Type species: Pseudoneoponera verecundae Donisthorpe, by original designation. Synonymy:
Wilson, 1958g: 361 (Pseudoneoponera junior synonym of Bothroponera)
;
Bolton, 1995b: 45 (Bothroponera junior synonym of Pachycondyla)
.
- OVERVIEW.
This large and diverse genus is found in the tropical and subtropical regions throughout the world. Only three species reach the United States. They nest in soil either in the open or under rocks and logs. Most species are general predators and scavengers. Some are specialized predators on termites, and one is known to harvest seeds (Brown, W.L. 2000: 60 in Agosti, Majer, et.al. Ants: standard methods for measuring and monitoring biodiversity). Queens are absent in some species with brood produced by fertilized workers. This large and complex genus has traditionally been a "dumping ground" for species that do not conveniently fit elsewhere. Shattuck is in the process of dividing this group (S. Shattuck, personal communication).
- DISTRIBUTION.
Southeastern USA.
- ZOOGEOGRAPHICAL REGIONS. Worldwide.

Photo of Pachycondyla stigma (Fabricius) from FL, Palm Beach Co.. Courtesy of Ant Web, U. of Calif. Davis
- RECOGNITION.
Length 7-13 mm. Mandible triangular with long masticatory margins containing many small teeth. Frontal carinae converge posteriorly; toward the front they flare out to form distinct lateral lobes that (in frontal view) conceal the antennal insertions. Eye large. Mid and hind tibiae with two apical spurs (the outer spur often nor more than half as long as the inner); one spur pectinate and one simple. (Shattuck, 1999. Australian Ants).
- TRIBE. Ponerini.
- REVISIONS.
Emery, 1890b: 7174.
- TAXONOMY.
Arnold, 1926: 202 (genus Hagensia)
.
Smith, M.R. 1943e: 282, 284 (description of m. (in part as Euponera (Brachyponera), and Neoponera (Neoponera))
.
Smith, M.R. 1947f: 536537, 539, pl. 2 fig. 8, pl. 3, fig. 11, 12 (in part as Neoponera (Neoponera), Euponera (Brachyponera), and Euponera (Trachymesopus))
.
Creighton, 1950a: 4244, pl. 5, fig. 14
(figure (lateral view) of w.q.m., wing, in part as Neoponera, Euponera (Brachyponera), and Euponera (Trachymesopus))
.
Arnold, 1951: 5356 (Afrotropical - out of date)
.
Arnold, 1952a: 460473 (Afrotropical - out of date)
.
Wilson, 1958g: 352, 362 (Trachymesopus not a subgenus of Euponera)
.
Kempf, 1960f: 423428 (New World species of Trachymesopus.)
.
Kempf, 1961e: 189204 (Brazilian species of Pachycondyla)
.
Brown, W.L. 1963a: 68
(three species groups of Trachymesopus)
.
Kempf, 1964e: 4952 (key to Neotropical species of Pachycondyla)
.
Brown, W.L. 1973b: 178185
(generic synonymy)
.
Bolton, 1994: figs. 467, 469, 495, 496 (SEM (full face and lateral view) of w.)
.
Xu, Z. 1994b: 181185 (China, in Chinese)
.Shattuck, 1999: 196198.Bolton, 2003: 44, 165, 280. - MISCELLANEOUS.
Hölldobler, B. & Wilson, 1990: 157, 234, 239, 241, 250251, 262, 263, 264, 268, 277, 279281, 292294, 347, 374376, 385, 394, 557 (trails, adult transport, colony founding, exocrine glands, foraging, larva, tandem running)
.
Orivel, Malherbe & Dejean. 2001: 449456 (adaption of pretarsus for arboreal living)
.
- ALL REFERENCES
Pachycondyla chinensis (Emery). The small colonies are found in moist, rotten wood or in soil beneath objects. They prefer dark, damp places. Food consists of small arthropods. Accidentally introduced into North America (Smith, D.R. 1979: 1341).
-
chinensis. Ponera nigrita subsp. chinensis
Emery, 1895m: 460 (w.)
. [China]
Wheeler, W.M. 1921c: 530 (q.)
. Combination in Euponera (Brachyponera): Emery, 1909d: 367; in Brachyponera: Brown, W.L. 1958h: 22; in Pachycondyla: Brown, W.L. in Bolton, 1995b: 304. Junior synonym of solitaria Smith, F., 1874b and hence first available replacement name: Brown, W.L. 1958h: 22.
-
solitaria. Ponera solitaria
Smith, F. 1874b: 404 (w.)
. [Japan, (Hiogo); BMNH] Combination in Euponera (Brachyponera): Emery, 1901b: 47. Junior primary homonym of Ponera solitaria Smith, F. 1860b: 103. Synonymy:
Bolton, 1995b: 309.
- DISTRIBUTION. VA, NC, GA; Japan, China, and adjacent areas.
- TAXONOMY.
Smith, M.R. 1934b: 384387
(description, first record for N. Amer.)
.
Smith, M.R. 1934d: 558, 559561, fig. 1
(description and figure (lateral view) of w., brief natural history, as Euponera (Brachyponera) solitaria)
.
Smith, M.R. 1947f: pl. 3, fig. 12 (figure (lateral view) of w. (as Euponera (Brachyponera) solitaria))
.
Creighton, 1950a: 4546
(as Euponera (Brachyponera) solitaria)
.
Wheeler, G.C. & Wheeler, J. 1952c: 629 (larva)
.
- ALL REFERENCES
Pachycondyla harpax (Fabricius). Colonies of about 150 individuals are found in rotten logs and stumps or in soil beneath objects. Workers avoid direct sunlight and forage in the morning and in shade for other insects and myriapods on which they feed. There are both ergatoid and normal females (Smith, D.R. 1979: 1340).
-
harpax. Formica harpax
Fabricius, 1804: 401
(w.)
. [South America, ("Meridional America"); No types in USA.]
Wheeler, W.M. 1900b: 4, fig. 2
(q.m.)
. Combination in Pachycondyla: Roger, 1863b: 18. Note that the correct page number for the first description of the queen and male is "Wheeler, W.M. 1900b: 4" not "Wheeler, W.M. 1900b: 19" as in Bolton, 1995b: 305.
-
montezumia. Pachycondyla montezumia
Smith, F. 1858a: 108 (q.m.)
. [Mexico]
Forel, 1899b: 12 (w.)
. Junior synonym of harpax: Roger, 1862c: 288; Roger, 1863b: 18; Dalla Torre, 1893: 34. Revived from synonymy as subspecies of harpax: Forel, 1899d: 12; Emery, 1911e: 75. Synonymy:
Brown, W.L. 1950g: 247
.
-
amplinoda. Pomera amplinoda
Buckley, S.B. 1866: 171 (w.)
. [USA, TX, (central)] Junior synonym of villosa: Forel, 1899b: 14. Synonymy:
Emery, 1911e: 75 (P. amplinoda junior synonym of P. montezumia)
;
Brown, W.L. 1950g: 247
(P. montezumia junior synonym of P. harpax)
.
-
orizabana. Pachycondyla orizabana
Norton, 1868c: 8 (w.)
. [Mexico] Junior synonym of montezumia: Emery, 1911e: 75. Synonymy:
Emery, 1911e: 75 (P. orizabana junior synonym of P. montezumia)
;
Brown, W.L. 1950g: 247
(P. montezumia junior synonym of P. harpax)
.
-
orizabana. Pachycondyla orizabana
Norton, 1868a: 64. Nomen nudum. Synonymy:
Bolton, 1995b: 308.
-
dibullana. Pachycondyla harpax var. dibullana
Forel, 1901k: 347 (w.)
. [Colombia]
Forel, 1912d: 39 (q)
. Synonymy:
Brown, W.L. 1950g: 247
.
-
irina. Pachycondyla harpax var. irina
Wheeler, W.M. 1925a: 5 (w.)
. [Guatemala] Synonymy:
Brown, W.L. 1950g: 247
.
-
concinna. Pachycondyla harpax var. concinna
Wheeler, W.M. 1925a: 5 (w.q.)
. [Brazil] Synonymy:
Brown, W.L. 1950g: 247
.
- DISTRIBUTION. LA, TX; south to Brazil; W. Indies.
- TAXONOMY.
Wheeler, W.M. 1900b: 46, fig. 2, 3
(figure (top view) w.q.m., lateral view of w.)
.
Wheeler, W.M. 1908h: 401403
(description w.q.m.)
.
Smith, M.R. 1943e: fig 3A (figure (lateral view) of petiole and gaster of m.)
.
Smith, M.R. 1947f: pl. 2, fig. 8 (figure (lateral view) of w.)
.
Brown, W.L. 1950g: 247248
(species synonymy)
.
Creighton, 1950a: 44, pl. 6, figs. 14
(figure (lateral view) of w.q.m., wing, as P. harpax subsp. montezumia)
.
Kempf, 1961e: 194.
- MISCELLANEOUS.
Wheeler, W.M. 1900b: 131
.
Haskins & Enzmann, 1938: 150151 (formation of new colony, in Canal Zone)
.
Cokendolpher & Francke, 1990: 15 (nest sites)
.
- ALL REFERENCES
Pachycondyla stigma (Fabricius). This species prefers to nest in moist, dead logs or stumps, occasionally under stones. It is apparently a tramp species distributed by commerce outside the New World. Whether or not it is endemic or adventive to Florida is unknown (Smith, D.R. 1979: 1340).
-
stigma. Formica stigma
Fabricius, 1804: 400
(q.)
. [South America]
Wheeler, G.C. & Wheeler, J. 1952c: 627 (l.)
. Combination in Ponera: Mayr, 1863a: 450; in Pachycondyla (Pseudoponera): Emery, 1900b: 314; Emery, 1901b: 46; in Euponera (Pseudoponera): Forel, 1901i: 398; in E. (Trachymesopus): Emery, 1911e: 85; in Trachymesopus: Wilson, 1958g: 355; Kempf, 1960f: 424; in Mesoponera: Wheeler, G.C. & Wheeler, J. 1976a: 55; in Pachycondyla: Brown, W.L. in Bolton, 1995b: 310. Current subspecies: nominal plus attrita, rufescens.
-
quadridentata. Ponera quadridentata
Smith, F. 1859a: 143 (q.)
. [Indonesia, (Aru Island)]
Roger, 1860: 285
(w.)
. Combination in Pachycondyla (Psdeudoponera): Emery, 1900b: 314; in Euponera (Trachymesopus): Emery, 1911e: 85. Variety / subspecies of stigma: Emery, 1897d: 552; Karavaiev, 1926d: 419. Synonymy:
Emery, 1887b: 434;
Wilson, 1958g: 355.
-
myopa. Ponera myopa
Roger, 1860: 285
(w.)
. [No type locality given] Synonymy:
Roger, 1860: 285
(myopa junior synonym of quadridentata)
;
Emery, 1887b: 434 (quadridentata junior synonym of stigma)
.
-
americana. Ponera americana
Mayr, G. 1862: 722 (w.)
. [Colombia] Synonymy:
Emery, 1887b: 434.
-
nixoni. Euponera (Trachymesopus) nixoni
Donisthorpe, 1943e: 441 (q.)
. [New Guinea]
Donisthorpe, 1948b: 299 (m.)
. Synonymy:
Wilson, 1958g: 355.
-
brunneus. Euponera (Trachymesopus) brunneus
Donisthorpe, 1948b: 300 (w.q.)
. [New Guinea] Synonymy:
Wilson, 1958g: 355.
-
sexdentatus. Euponera (Trachymesopus) sexdentatus
Donisthorpe, 1949c: 746 (w.)
. [New Guinea] Synonymy:
Wilson, 1958g: 355.
- DISTRIBUTION. FL and AL; W. Indies, Mexico south to northern Argentina; southern China to Samoa, northern Queensland.
- TAXONOMY.
Smith, M.R. 1934d: 558, 561564
(description w. (as Euponera (Trachymesopus) stigma))
.
Smith, M.R. 1937: 824824
(natural history in Puerto Rico, as Euponera (T.) stigma)
.
Smith, M.R. 1947f: 534 (figure of worker (as Neoponera (Neoponera) villosa))
.
Creighton, 1950a: 46, pl. 7, figs. 14
(figure (lateral view) of w.q.m., wing, (as Euponera (Trachymesopus) stigma))
.
Wilson & Taylor, 1967b: 22 (Polynesia)
.
Wheeler, G.C. & Wheeler, J. 1976a: 4164 (descriptions of larva from different localities, as Mesoponera stigma)
.
- ALL REFERENCES
- PHOTOS.
Pachycondyla villosa (Fabricius). Colonies occur in the soil and in logs and stumps. Workers run rapidly in the bright sun in search of insects on which they feed. They can sting severely. This is the largest ponerine ant in the USA (Smith, D.R. 1979: 1340).
-
villosa. Formica villosa
Fabricius, 1804: 409
(w.)
. [Central America, ("Meridional America"); No types in USA.]
Roger, 1861a: 2
(w.q.)
. Combination in Pachycondyla: Mayr, 1862: 720; in Neoponera: Emery, 1901b: 43; in Pachycondyla: Brown, W.L. in Bolton, 1995b: 311. Current subspecies: nominal plus inversa.
-
bicolor. Ponera bicolor
Guérin-Méneville, 1844a: 424 (w.)
. [Mexico] Synonymy:
Roger, 1861a: 1
;
Roger, 1862c: 288
.
-
pilosa. Ponera pilosa
Smith, F. 1858a: 95 (m.)
. [Brazil] Synonymy:
Roger, 1861a: 1
;
Roger, 1862c: 288
.
-
pedunculata. Ponera pedunculata
Smith, F. 1858a: 96, pl. 6, fig. 25 (w.)
. [Brazil] Synonymy:
Roger, 1861a: 1
;
Roger, 1862c: 288
.
- DISTRIBUTION. Southern Texas (as far north as San Antonio); south to northern Argentina.
- TAXONOMY.
Wheeler, W.M. 1908h: 404
.
Gallardo, 1918c: 56.
Smith, M.R. 1947f: pl. 3, fig. 11 (figure (lateral view) of w. as Neoponera (Neoponera) villosa)
.
Creighton, 1950a: 43, pl. 5, figs. 14
(figure (lateral view) of w.q.m., wing, as Neoponera villosa)
.
Wheeler, G.C. & Wheeler, J. 1971b: 1205 (larva)
.
- MISCELLANEOUS.
Cokendolpher & Francke, 1990: 16 (occurrence in TX)
.
- ALL REFERENCES
Genus Pachycondyla
URL: http://www.cs.unc.edu/~hedlund/playpen/dev/ants/catalog/
Last updated: Sat May 19 09:43:12 EDT 2007
Copyright 2006, Kye S. Hedlund, University of North Carolina, hedlund@cs.unc.edu